WillowTree Ventures, also referred to as WillowTree Investments, is an early-stage venture capital firm founded in 2020. Headquartered in the United States, with locations in New York City and Rye, the firm focuses on technology-driven companies with significant innovation and growth potential. They adopt an "AI-first" investment thesis, targeting sectors such as artificial intelligence, enterprise technology, fintech, vertical SaaS, e-commerce, and sustainability. WillowTree Ventures typically co-invests up to $500,000 in funding rounds ranging from pre-seed to Series A. The firm emphasizes investing in experienced entrepreneurs at the concept stage, often prior to revenue or a full team, based on their credible vision. They strategically co-invest rather than leading funding rounds, focusing on ventures that drive the adoption and operationalization of generative AI in enterprises, while avoiding direct investments in Large Language Models (LLMs).